| Adaptation & Selection |
Variation: If all members of a species were identical there could be no evolution; which organisms survive and reproduce would be determined solely by chance and would not result in any changes over time. But
large amounts of empirical data indicate that organisms of the same
species are not genetically identical. For example, an analysis
of human DNA reveals that when we compare two unrelated individuals,
we can expect one sequence difference for every 1200 base pairs
we examine. Since each human cell has approximately
6,000,000,000 base pairs of DNA, that is a lot of differences (more here).
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If we were to measure a large enough population of organisms with respect to one of these "quantifiable" traits, we would determine the frequency distribution for the trait. For many traits, this distribution is a normal distribution. You can identify a normal distribution because it is symmetrical around a peak; the peak value is generally the mean of the distribution. Generally traits that fall into a normal distribution are determined by the action of multiple genes. |
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On the other hand some traits, such as blood type (that is, whether you have O, A, B or AB blood type), are discontinuous. Such traits are generally controlled by variations in one gene. Not all traits effect the ability of an organism to survive or reproduce successfully. Natural selection acts directly on traits that alter reproductive success. At the same time, it is possible for two traits to be linked so that selection of one leads to selection of the other. The genes involved can be located near one another on a particular chromosome or it may be that the process of producing the selected trait also produces the other "non-selected" trait. If this is the case, then the non-selected trait will be found in the population not because it is in and of itself advantageous, but because it necessarily accompanies the selected trait. This is worth keeping in mind whenever someone tries to explain the presence of a particular trait based on its presumed benefits. |
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Directed selection: What happens when certain values of a trait positively impact survival while values at the other extreme negatively impact survival? Assuming that the trait has a genetic component, over time the mean will move in the direction of the positive effect, and away from the negative effect. The shape of the distribution may change depending on a number of factors. |
In
the
case above |
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Conservative selection: Another possibility is that individuals with traits at either end of the population distribution are at a reproductive disadvantage compared with organisms that are closer to the mean. With each generation, the contribution of these "outliers" to the next generation will be reduced. The distribution mean will remain constant, but the standard deviation will decrease. The stronger the disadvantage the outliers face, the narrower the distribution will become. In the end, the size of the standard deviation will reflect the rate at which new variation enters the population. |
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Disruptive selection: A third possibility is that the organisms with traits at the extremes of the population distribution actually have a reproductive advantage over those nearer the mean. This type of behavior would tend to expand the population distribution, while mating between organisms at the extremes would tend to bring the population distribution back to the original state. If organisms at the extremes preferred to mate with organisms with the same form of the trait, the population would split into two. |
| Such a mating preference could
itself be a selectable trait, and would represent the formation of
a reproductive barrier. |
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Use Wikipedia |
revised
25-Aug-2008
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