| Symbiosis: mitochondria and chloroplasts |
In aerobic respiration, high energy electrons derived from food molecules (such as carbohydrates) are delivered to O2 forming water; the energy released is used primarily to generate ATP. In oxygenic photosynthesis electrons, derived from water, are energized by light, and that energy is used to generate ATP and carbohydrates. |
In aerobic bacteria, electron transport chains are localized in the plasma membrane; in cyanobacteria, the photosystems are located on internal membrane systems. The same mechanisms (aerobic respiration and photosynthesis) occur within eukaryotic cells, not on the plasma membrane, but within distinct cytoplasmic organelles: mitochondria for aerobic respiration and chloroplasts for photosynthesis. |
| Based on large amounts of molecular data, it is clear that the mitochondria of eukaryotes were derived from α-proteobacteria-like bacteria, while chloroplasts were derived from cyanobacteria-like bacteria. |
The exact details of the endosymbiotic process, however, remain unclear. In particular, the exact nature of the original host cell remains the subject of sometimes heated debate. |
In part, this is due to the large amount of gene transfer from the bacterial endosymbiont to the host cell nucleus . At the same time, it is not clear exactly where the eukaryotic nucleus came from. Did the original endosymbiotic event involve more than two types of organisms? |
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The original endosymbiotic events that produce modern eukaryotes occurred approximately 2 billion years ago, making unambiguous details somewhat difficult to come by. What does appear to be the case, is that all eukaryotes may well be descended from a single endosymbiotic event. Plants appear to be descended from a second endosymbiotic event. |
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revised
25-Aug-2008
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